Staff+ Application Security Engineer

<div class="content-intro"><h2><strong>About Anthropic</strong></h2> <p>Anthropic’s mission is to create reliable, interpretable, and steerable AI systems. We want AI to be safe and beneficial for our users and for society as a whole. Our team is a quickly growing group of committed researchers, engineers, policy experts, and business leaders working together to build beneficial AI systems.</p></div><div> <h2>About the role:</h2> </div> <div> <p>Anthropic's Application Security team secures the systems that build, serve, and increasingly <em>are</em> Claude. The attack surface is unlike most AppSec work: multi-agent orchestration, sandboxed code execution, agents holding delegated credentials, untrusted tool output crossing trust boundaries — problems with little prior art and no off-the-shelf playbook.</p> <p>The way the team works is also different. We use Claude as our primary tool across every part of the job: it drives our static analysis, drafts and fixes vulnerabilities as pull requests, performs first-line bug bounty triage, and assists threat modeling for design reviews. The human work is the judgment layer — system-level reasoning, deciding what matters, and building the next thing the model can't do yet.</p> <p>This is a builder's role on a senior team. We hire engineers who ship production systems <em>and</em> clear a hands-on threat-modeling bar — people who can find the vulnerability but would rather build the system that finds them all. Every engineer owns a system end-to-end, and the team's work has shaped customer-facing product security — including Claude Code's security review tooling, its security guidance plugin, sandboxing, and auto mode.</p> <h2>Key responsibilities:</h2> <ul> <li>Design, build, and operate Claude-powered security systems — LLM-driven code analysis, automated vulnerability remediation, AI-assisted threat modeling — and own one or more of them end-to-end, including the cross-functional relationships that come with it</li> <li>L